Index
Practical ecological knowledge for the temperate reader.

Campanula Sp. - Bellflower

Family: Campanulaceae (Bellflower family) [E-flora]

"Annual, perennial herb, from taproot, fibrous roots, or rhizome, hairs 0 to dense. Stem: reclining or erect, branched, 2–60[150] cm, 4-angled. Leaf: basal or not, cauline, generally lanceolate to ovate, thin, fleshy, or leathery, entire to toothed, sessile or petioled, gradually reduced distally to bracts in inflorescence. Inflorescence: raceme and/or flowers 1 [head, spike, panicle]; terminal or axillary. Flower: not inverted; corolla cylindric to funnel- or bell-shaped, white to deep blue, cut 1/4–2/3 [to all the way] to base, lobes narrow- to wide-triangular; ovary inferior, hemispheric, spheric, or oblong to obconic. Fruit: open by 2–3 lateral pores. Seed: 0.6–3.5 mm, oblong or fusiform.
± 400 species: northern hemisphere; many cultivated, some medicinal. (Latin: little bell, from corolla shape) [Roquet et al. 2008 Syst Bot 33:203–217] Campanula prenanthoides moved to Asyneuma." [Jepson]

Local Species;

"The genus Campanula comprises approximately 300 species distributed across the Northern Hemisphere (Cook 1951), many of these in mountainous areas. The genus generally inhabits meadow and subalpine regions, many species requiring full sun for optimal development. All species are herbaceous, and the name refers to the blue, bell-shaped flowers of the majority of the species. They are perennials, biannuals, or annuals. The growth pattern is characterized by a distinct vegetative phase, often with development of rosettes, and a generative phase, induced by long photoperiods, vernalization, or both, where the rosettes develop into multi- or single-flowering inflorescences. Usually, a flowering shoot dies after seed maturation. In biannual and annual species each seedling usually forms one rosette. The perennial species develop sideshoots or more or less underground runners each of which becomes a rosette during the following growth season." [Brandt.,1997]

"About three hundred species of Campanula occur in the northern temperate region. West European species such as C. poscharskyana, C. portenschlagiana, and C. glomerata have been grown in flower gardens since the 16th century or, in the case of C. medium, since the Medieval period or earlier. More recent introductions include C. barbata, C. isophylla (Falling Stars), and C. punctata, and smaller species such as C. carpatica and C. cochlearifolia. Although introduced only in the 1980s, the Korean C. takesimana has rapidly become widely grown. Flowers are commonly bell-shaped, but may have a flatter, star shape, and may be borne on tufted, sprawling, or tall erect stems." [Prance TCHP]

  1. Campanula lasiocarpa - Mountain harebell (Haida Gwaii) [E-flora]
  2. Campanula medium - Canterbury-bells [E-flora]
  3. Campanula persicifolia - peach-leaved bellflower [E-flora]
  4. Campanula rapunculoides - creeping bellflower [E-flora]
  5. Campanula rotundifolia - common harebell [E-flora]
  6. Campanula scouleri - Scouler's harebell [E-flora]
  7. Campanula trachelium - nettle-leaved bellflower [E-flora]

Medicinal Uses of Various sp.

Campanula spp.; "Infusion of eastern species used by Native Americans to treat whooping cough and tuberculosis." [Meuninck EWPUH]

"A few species, Campanula glomerata, C. persicifolia, C. rotundifolia, C. bononiensis, C. sibirica, and C. patula have been used locally for preparation of traditional drugs in Russian folk medicine (Barnaulov et al. 1983, 1984 and references therein), and, in Italy, similar use has been made of C. medium, C. cervicaria, C. rotundifolia, C. latifolia, and C. trachelium (Gastaldo 1978). Preparations were used to treat epilepsy, nervous diseases, coughs, headache, rheumatism, and inflammation. Many of the medicinal qualities attributed to these species are similar to the use in oriental medicine of drugs made from the closely related Platycodon grandifiorum (Ozaki 1995; Tada et al. 1995a), a species originally classified in the genus Campanuia, and from species of the allied genera Adenophora (Kuang et al. 1991) and Codonopsis (Wang et al. 1995)." [Bajaj MAPS 10]

Major Constituents and Therapeutic Values of Chinese Medicinal Herbs [CRNAH]

Scientific Name

Common Chinese and (English) Name

'''Major Constituents and (sources)'''

Therapeutic Values*

Campanula gentianoides Lam. C. giauca Thunb. C. grandiflora Jacq. (Syn.Platycodon grandiflorum)

Jie Geng

(root) "Saponins, inulin, platycodigenin."

As an expectorant

Campanula glomerata; C. glomerata L. var. dahurica; C. punctata Lam

Feng Lin Cao

(whole plant) "Quercetin, isorhamnetin, kaempferol, hyperoside, isoquercetin, trifolin, chlorogenic acid, methyl caffeate, coumaroylquinic acid."

For throat infection, headache

Chinese and North American Medicinal Herbs Belonging to the Same Genus and Different Species: Major Constituents and Therapeutic Values

Source

Scientific Name

Major Constituents

Therapeutic Values*

N.A.

Campanula rotundifolia L.; C. palustris L.

"Lutein-7-primveroside, luteolin-7-0- beta-D-glucopyranosil, rhamnetin-3-0- beta-D-galactoside, esculetin, caffeic, n-coumaric, ferulic acids."

"For faintness or a weak heart, stop bleeding, reduce swelling."

Campanula parryi A. Gray. Bluebell. - "The blossoms are chewed and the saliva is applied to the skin to render it depilous. According to the statements of the Zuñi this medicine removes only young hair. It belongs to the people in common. For reducing bruises the chewed root is applied with bandages. When employed for this purpose the remedy belongs only to the fraternities." [Stevenson Zuni]

Food Uses of various Campanula Sp.

"Wild vegetables for salads are harvested especially during the spring season. The local people prepare mixtures with up to ten species: Campanula, Cichorium, Crepis, Hypochoeris, Reichardia, Sonchus and Valerianella representing the most used genera. The local people also ascribe depurative and diuretic action to such food." [Andrea Pieroni,1999]

Names Range Uses
Campanula fenestrellata - Adriatic Bellflower S. Europe - Adriatic Coast. Leaves - raw or cooked[K]. A pleasant sweet flavour[K]. It makes a very acceptable addition to mixed salads in the winter[K]. Flowers - raw. Produced in abundance, they have a pleasant sweet flavour and make a decorative addition to the salad bowl[K].(1)
Campanula garganica - Adriatic Bellflower S. Europe - Italy. Leaves - raw or cooked. The leaves are quite small but have a pleasant mild flavour, and make a very acceptable addition to salads[K]. Flowers - raw or cooked. Slightly sweet[K].(2)
Campanula glomerata - Clustered Bellflower, Dane's blood, Clustered Bellflower Most of Europe, including Britain, to temperate Asia. Leaves - raw or cooked. A mild flavour with a pleasant sweetness, it can be used as a major ingredient in salads[K]. Flowers - raw[K]. Beautiful to look at, they have a pleasant sweetness and make a very attractive decoration to a salad[K].(3)
Accessed Feb 10, 2015

Other Uses

Decorative

Campanula patula L. & Campanula persicifolia L. Aerial part - Ornamental: bouquet elaboration [Gras et al.,2016] Campanula glomerata L. & Campanula persicifolia L. cultivated and sold in pots as ornimentals [Kasper-Pakosz et al.,2016]

Phytochemicals

p-Hydroxybenzoic Acid: "Most of the 16 anthocyanins that are acylated with p-hydroxybenzoic acid (Appendix A) have been isolated from flowers of Delphinium hybridum,109,119,183 Aconitum chinense,184 Consolida armeniaca (Ranunculaceae),115 Campanula species (Campanulaceae),117,185,186 or roots of Ipomoea batatas (Convolvulaceae).187,188 In the examined species from Ranunculacea and Campanula, one to as many as four p-hydroxybenzoyl moieties belong to the anthocyanidin 7-glycoside." [Andersen FCBA]

"The five major anthocyanins found in the Campanula species examined so far are shown in Fig. 12. Campanin, rubrocampanin, and monodeacyl- campanin are known only from Campanula (Terahara et al. 1990; Brandt et al. 1993). Bisdeacylplatyconin was described first as a degradation product of platyconin from Platycodon grandifiorum (Goto et al. 1983) and violdelphin was identified in Delphinium hybridum (Kondo et al. 1990). Both were later found to occur also in Campanula (Brandt et al. 1993). Genotypes containing rubrocampanin as the major anthocyanin have pink flowers (Terahara et al. 1990), and mutants where the biosynthesis is blocked after bisdeacyl- plathyconin have a very pale, light purple color (Brandt 1990). Flowers of chemotypes with the acylated delphinidin derivatives campanin, mon- odeacylcampanin, and violdelphin all have purple-blue colors with the same hue; they cannot be distinguished without a chemical analysis (Brandt 1990; K. Brandt, unpubl.)." [Bajaj MAPS 10]

[Bajaj MAPS 10]

Cultivation & Propagation

"Traditional propagation is either by seed or cuttings; biannual species, such as Campanula medium and C. pyramidalis, are seed-propagated, while the perennial species may be vegetatively propagated by cuttings or runners during the vegetative phase. If the plants enter the generative phase, the shoots rapidly lose the ability to form roots (Moe and Andersen 1988). [Brandt.,1997]

"Mountain natives, bellflowers prefer full sun to light shade; well-drained, organic soil; and regular watering. Serbian bellflower. C. poseharskyana. tolerates drought, but most other bellflowers do not. A layer of mulch helps keep the root zone cool and holds in moisture during hot summer weather: a win­ ter mulch protects roots from heaving. Divide plants in spring every 3-4 years." [HNIDC Ellis]

"For floriculture and ornamental crops, regulation of photoperiodic response is crucial and, for that reason, foundational knowledge of species-specific photoperiod and temperature requirements for flowering is now available at least for the major ornamental species. For example, use of night interruption lighting to promote flowering of long-day plants (such as campanula) or to prevent flowering of short-day plants (such as chrysanthemum) is widely practiced." [Kozai PF]


Campanula rapunculus - Rampion

Campanulaceae - Campanula rapunculus-5 [1]
Campanula rapunculus

"The native range of this species is Europe to Medit. and Iran. It is a biennial and grows primarily in the temperate biome." [PWO]

"Rampion (Campanula rapunculus). The tubers resemble tiny potatoes and were once popular in salads and as a table vegetable. It is still cultivated to some extent in France, Germany, and Switzerland. Historically, it was grown as a garden vegetable even during Roman times, with the center of cultivation in the Rhine Valley. Its Latin name means little turnip, and it was evidently an important food among the Gauls and early Germans, perhaps with some now lost sacred connections. Numerous folk legends survive dealing with personifications of this tuber, including the famous Grimm fairy tale about Rapunzel, and the story of the first king and queen of ancient Poland—the queen’s name was rampion." [Katz EFC]

"C. rapunculus Linn. RAMPION. Europe, Orient, north Africa and northern Asia. This biennial plant was formerly much cultivated in gardens for its roots as well as its leaves. Loudon says the latter are excellent, eaten raw as a salad or boiled as a spinach, and the root, which has the flavor of walnuts, is also eaten raw like a radish or mixed with salads, either raw or boiled and cold. It is much cultivated in France and Italy, says Johns." [Sturtevant EWP]

"Figure 4.6 also shows that 184 species (70 % of the wild vegetables) are consumed raw. Those species considered of better quality because of their pleasant flavour are used in salads. Besides some aforementioned species, the small leaves and stems of Montia fontana, the young shoots of Chamaerops humilis, the basal leaves of Reichardia picroides (L.) Roth and the leaves and roots of Campanula rapunculus L. are some of the species most widely consumed raw in salads." [Tardio MWEP]

[Freitas,2004]

References

Image References


Campanula lasiocarpa - Mountain harebell (Haida Gwaii)

Campanula lasiocarpa 04 [1]
Campanula lasiocarpa

Image References

"Campanula lasiocarpa is a PERENNIAL growing to 0.2 m (0ft 8in).
It is hardy to zone (UK) 4. It is in flower in July, and the seeds ripen from Aug to September. The flowers are hermaphrodite (have both male and female organs) and are pollinated by Bees, flies, beetles, lepidoptera, self.The plant is self-fertile.
Suitable for: light (sandy) soils, prefers well-drained soil and can grow in nutritionally poor soil. Suitable pH: acid, neutral and basic (alkaline) soils. It cannot grow in the shade. It prefers moist soil." [PFAF]

"General: Perennial herb from a thin, branching rhizome; stems single to several, erect or ascending, 2-20 cm tall, smooth or slightly hairy." [IFBC-E-flora]

"Habitat / Range Moist to mesic meadows, forests and rocky slopes in the montane to alpine zones; common N of 52degreeN; amphiberingian, N to AK, YT, and NT, E to W AB and S to N WA; E. Asia." [IFBC-E-flora]

"Gravelly and sandy slopes in C. and N. Japan[58]. Crevices in rocks and on rocky slopes in alpine areas at elevations of 1600 - 3100 metres on Honshu, 1000 - 2000 metres on Hokkaido[275]." "E. Asia - China, Japan to Western N. America - Rocky Mountains." [PFAF]

Origin Status: Native [E-flora]

Edible Uses

Propagation "Seed - surface sow spring in a cold frame. The seed usually germinates in 2 - 4 weeks at 18oc. Easy[221]. When they are large enough to handle, prick the seedlings out into individual pots and grow them on in a cold frame for at least their first winter. Plant them out into their permanent positions in late spring or early summer, after the last expected frosts. Basal cuttings in spring. Harvest the shoots when they are about 10 - 15cm long with plenty of underground stem. Pot them up into individual pots and keep them in light shade in a cold frame or greenhouse until they are rooting well. Plant them out in the summer. Division in spring or autumn. Very easy, larger clumps can be replanted direct into their permanent positions, though it is best to pot up smaller clumps and grow them on in a cold frame until they are rooting well. Plant them out in the summer or following spring." [PFAF]

Cultivation "Prefers a moist but well-drained rich sandy loam and a neutral or alkaline soil in sun or partial shade[1, 200]. A rock garden plant, it is difficult to grow in the garden needing a very gritty soil and perfect drainage[1]. It grows freely amongst rocks or in a loose scree, but is very impatient of winter wet[221]. Plants are hardy to at least -15oc[200]. Slugs are very partial to this plant[221]. Members of this genus are rarely if ever troubled by browsing deer[233]. The species in this genus do not often hybridize and so seed can generally be relied upon to come true[221]. The plants are self-fertile[221]. Plants produce seed freely in British gardens[221]. There is at least one named variety, selected for its ornamental value. 'Alba' has white flowers[200]." [PFAF]

Synonyms
Campanula lasiocarpa subsp. latisepala (Hultén) Hultén [E-flora]
Campanula latisepala Hultén [E-flora]
Campanula latisepala var. dubia Hultén [E-flora]

References


Campanula medium - Canterbury-bells

Campanulaceae - Campanula medium-4 [1]
Campanula medium

Image References

"Campanula medium is an evergreen Biennial growing to 1.2 m (4ft) by 0.3 m (1ft in) at a medium rate.
It is hardy to zone (UK) 8. It is in leaf 12-Jan It is in flower in July, and the seeds ripen from Aug to September. The flowers are hermaphrodite (have both male and female organs) and are pollinated by Bees, flies, beetles, lepidoptera, self.The plant is self-fertile.
Suitable for: light (sandy) and medium (loamy) soils and prefers well-drained soil. Suitable pH: neutral and basic (alkaline) soils and can grow in very alkaline soils.
It can grow in semi-shade (light woodland) or no shade. It prefers moist soil." [PFAF]

"General: Biennial herb from a fibrous root or rhizome; stems single to several, erect or ascending, 35-90 cm tall, bristly-hairy." [IFBC-E-flora]

"Habitat / Range Mesic disturbed areas in the lowland zone; rare garden escape in coastal SW BC; introduced from Europe." [IFBC-E-flora]

Status: Exotic [E-flora]

Edible Uses

Other Uses

Phytochemicals

"Campanula medium L. is a common ornamental that varies in color from white to pink to violet-blue flowers, originating in Southern Europe. The nine chemical structures of acylated pelargonidin, cyanidin, and delphinidin derivatives in the flowers of some Campanula species, including C. medium, have previously been studied (Brandt et al., 1993, Terahara et al., 1990, Toki et al., 2006, Toki et al., 2009)." [Tatsuzawa.,2023]

(-)-Lobeline (piperidine) - Lobelia hactleri, L. inJata, L. nocitianaefolia, L. tupa (S. Am. "Indian tobacco"), Campanula medium [seed] (Campanulaceae) - nACh-R agonist - a4B2 [4nM] (a7 antagonist) [anti-smoking use; racemate (Lobelidine) analeptic [Polya BTPBC]

Propagation
"Seed - best sown in an outdoor seedbed in May, the seedlings are transplanted when large enough and then placed in their final positions in the autumn[221]. The seed usually germinates in 2 - 4 weeks at 18oc[138]." [PFAF]

Cultivation
"Succeeds in most fertile well-drained soils[233], though it prefers a moist but well-drained rich sandy loam and a neutral or alkaline soil in sun or partial shade[1, 200]. A very ornamental plant, it is often grown in the flower garden and there are many named varieties[17]. The species in this genus do not often hybridize and so seed can generally be relied upon to come true[221]. The plants are self-fertile[221]. Plants usually self-sow in British gardens[221]. Members of this genus are rarely if ever troubled by browsing deer[233]." [PFAF]

"Short-long-day plants (SLDPs) flower only after a sequence of short days followed by long days. SLDPs, such as Trifolium repens (white clover), Campanula medium (Canterbury bells), and Echeveria harmsii (echeveria), flower in the early spring in response to lengthening days." [Zeiger PP]

References


Campanula persicifolia - peach-leaved bellflower

Campanula persicifolia 2015-06-20 3402 [1]
Campanula persicifolia

Image References

"Campanula persicifolia is an evergreen Perennial growing to 1 m (3ft 3in) by 0.5 m (1ft 8in).
It is hardy to zone (UK) 3 and is not frost tender. It is in leaf 12-Jan It is in flower from Jun to August, and the seeds ripen from Aug to October. The flowers are hermaphrodite (have both male and female organs) and are pollinated by Bees, flies, beetles, lepidoptera, self.The plant is self-fertile.
Suitable for: light (sandy) and medium (loamy) soils and prefers well-drained soil. Suitable pH: neutral and basic (alkaline) soils. It can grow in semi-shade (light woodland) or no shade. It prefers moist soil." [PFAF]

Origin Status: Exotic [E-flora]

"General: Perennial herb from a branching rhizome; stems single to several, erect, leafy, 35-90 cm tall, glabrous or sparsely hairy." [IFBC-E-flora]

"Habitat / Range Mesic disturbed areas in the lowland zone; rare garden escape in coastal SW BC; introduced from Eurasia." [IFBC-E-flora]

"Commons and open woods[13, 17]." "Europe to western and northern Asia. Locally naturalized in Britain." [PFAF]

Edible Uses

"C. persicifolia Linn. PEACH BELLS. Europe and north Asia. This plant has been used as food in England but has long since fallen into disuse. In France it is called cloche and is grown as a flowering plant." [Sturtevant EWP]

Propagation
"Seed - surface sow spring in a cold frame. The seed usually germinates in 2 - 4 weeks at 18°c[138]. When they are large enough to handle, prick the seedlings out into individual pots and grow them on in a cold frame for at least their first winter. Plant them out into their permanent positions in late spring or early summer, after the last expected frosts. Basal cuttings in spring[1]. Harvest the shoots when they are about 10 - 15cm long with plenty of underground stem. Pot them up into individual pots and keep them in light shade in a cold frame or greenhouse until they are rooting well. Plant them out in the summer. Division in spring or autumn[200]. Very easy[221], larger clumps can be replanted direct into their permanent positions, though it is best to pot up smaller clumps and grow them on in a cold frame until they are rooting well. Plant them out in the summer or following spring." [PFAF]

Cultivation
"Easily grown in ordinary garden soil[187]. Prefers a moist but well-drained rich sandy loam and a neutral or alkaline soil in sun or partial shade[1, 200]. Succeeds in light woodland[200]. Plants are hardy to at least -15°c[200]. The species in this genus do not often hybridize and so seed can generally be relied upon to come true[221]. The plants are self-fertile[221]. A very ornamental plant, there are many named varieties[187]. It was at one time grown as a culinary vegetable, but is now only grown as an ornamental plant[4]. The sub-species C. persicifolia crystalocalyx has larger leaves than the species and so is more suitable as a food crop[K]. A very long-lived and easily grown plant[187], it is best divided every other year[111]. Slugs are very fond of this plant and can cause severe damage even to large plants[K]. Members of this genus are rarely if ever troubled by browsing deer[233]. This plant is a potential winter salad plant, it retains a basal rosette of leaves all winter[K]." [PFAF]

Synonyms

References


Campanula rapunculoides - creeping bellflower

Campanula rapunculoides - Creeping Bellflower - Elmacık 3 [1]
Campanula rapunculoides

Image References

"Campanula rapunculoides is a PERENNIAL growing to 1.2 m (4ft) by 1 m (3ft 3in) at a fast rate.
It is hardy to zone (UK) 3. It is in flower from Jul to September, and the seeds ripen from Aug to October. The flowers are hermaphrodite (have both male and female organs) and are pollinated by Bees, flies, beetles, lepidoptera, self.The plant is self-fertile.
Suitable for: light (sandy) and medium (loamy) soils and prefers well-drained soil. Suitable pH: neutral and basic (alkaline) soils. It can grow in semi-shade (light woodland) or no shade. It prefers moist soil." [PFAF]

Origin Status: Exotic [E-flora]

General: Perennial or sometimes biennial herb from a branching rhizome; stems single to several, erect, over 50 cm tall, glabrous or sparsely hairy." [IFBC-E-flora]

"Habitat / Range Disturbed areas in the lowland and lower montane zones; frequent, scattered across extreme S BC; introduced from Eurasia." [IFBC-E-flora]

"Fields and woods[100]. Naturalised in Britain where it grows in fields and more or less disturbed grassy areas such as railway banks, occasionally in woods, usually near to houses[17]." "Europe, north to 65oN., east to W. Asia and the Caucasus. Introduced in Britain." [PFAF]

Edible Uses

"C. rapunculoides Linn. CREEPING BELLFLOWER. Europe and temperate Asia. This plant may be substituted in cultivation for rampion. It has long since fallen into disuse." [Sturtevant EWP]

Medicinal Uses

Lore

"RAMPION BELLFLOWER
(Campanula rapunculus) Not a British native, but established here and there, usually as an escape. The specific name, rapunculus, means a little turnip, and the roots are quite edible, either raw, or sliced in salads, or cooked, when they taste rather like parsnips." [DPL Watts]

Propagation
"Seed - surface sow spring in a cold frame. The seed usually germinates in 2 - 4 weeks at 18oc[138]. Easy[221]. When they are large enough to handle, prick the seedlings out into individual pots and grow them on in a cold frame for at least their first winter. Plant them out into their permanent positions in late spring or early summer, after the last expected frosts. Basal cuttings in spring[200]. Harvest the shoots when they are about 10 - 15cm long with plenty of underground stem. Pot them up into individual pots and keep them in light shade in a cold frame or greenhouse until they are rooting well. Plant them out in the summer. Division in spring or autumn[200]. Very easy, any part of the root will produce a new plant[221]." [PFAF]

Cultivation
"An easily grown plant succeeding in almost any soil[221], though it prefers a moist but well-drained rich sandy loam and a neutral or alkaline soil in sun or partial shade[1, 200]. It is slower growing and less spreading when grown in heavier soils[271]. Plants are hardy to at least -15oc[200]. The species in this genus do not often hybridize and so seed can generally be relied upon to come true[221]. The plants are self-fertile[221]. A beautiful plant, it was at one time cultivated as a culinary plant but has fallen into disuse[4]. The plant produces a mass of thick white roots which can spread at an alarming rate, especially in light soils. It can often fill an entire bed with its tenaceous roots, killing off less vigorous plants[271]. It is best grown in the wild garden where it can be allowed to romp without harm[221, 271]. Plants can also succeed when growing in thin grass[233]. Plants produce seed freely and often self-sow[221]. Slugs are very attracted to this plant, we have had great problems growing it on our Cornish trial grounds because the slugs eat out all the new shoots in spring and can kill even well-established specimens[K]. Members of this genus are rarely if ever troubled by browsing deer[233]." [PFAF]

"Weedy flowers such as creeping bellflower (Campanula rapunculoides) can quickly spread and overtake a garden, crowding out desirable plants." [Steiner NCG]

Synonyms

References


Campanula rotundifolia - Common harebell

Campanula rotundifolia Linné. – Campanule à feuilles rondes. – (Bluebell). [1]
Campanula rotundifolia

Image References

"Campanula rotundifolia is a PERENNIAL growing to 0.4 m (1ft 4in) at a medium rate. It is hardy to zone (UK) 3 and is not frost tender. It is in flower from Jul to September, and the seeds ripen from Aug to October. The flowers are hermaphrodite (have both male and female organs) and are pollinated by Bees, flies, beetles, lepidoptera, self.The plant is self-fertile.''' Suitable for: light (sandy) and medium (loamy) soils, prefers well-drained soil and can grow in nutritionally poor soil. Suitable pH: neutral and basic (alkaline) soils. It can grow in semi-shade (light woodland) or no shade. It prefers dry or moist soil." [PFAF]

"General: Perennial herb from a taproot, slender rhizomes or branched stem-base; stems single to several, erect or ascending, 10-80 cm tall, more or less glabrous." [IFBC-E-flora]

"Notes: There is little evidence to suggest that C. alaskana is anything other than a coastal race of the highly variable C. rotundifolia. Many taxonomists (e.g., Shetler 1963, Welsh 1974, Scoggan 1979, Kartesz 1994) do not recognize any infraspecific taxa within this species." [IFBC-E-flora]

Origin Status: Native [E-flora]

Habitat / Range

"Moist to dry meadows and rock outcrops in all vegetation zones; common throughout BC; circumboreal, N to AK, YT and NT, E to NF and S to TX, NM and N CA." [IFBC-E-flora]
"Dry grassy places and on fixed dunes, often in poor shallow soils throughout most of Britain[17]." "Northern Temperate regions of the world, including Britain, to latitude 70o N." [PFAF]

Edible Uses

Medicinal Uses

"The majority (59%) of native South Tyrolean medicinal plants remain understudied by medicine and pharmacology. However, considering the success rate (> 70%) of previously investigated plants (111) for which the traditional uses have been validated (79), we get an idea of the large unlocked phytotherapeutic and economic potential of as yet unexplored plants. In particular, the healing properties of the following largely unexplored plants should be analyzed, i.e., the highly quoted species Peucedanum ostruthium (wound healing, anti-inflammatory, digestive, and respiratory properties), ... Campanula rotundifolia (borreliosis), .... [Petelka,Joshua,et al.,2015]

Phytochemicals

Propagation
"Seed - surface sow in spring in a cold frame. Three or four weeks pre-chilling of the seed improves the germination rate[138]. The seed usually germinates in 2 - 4 weeks at 18oc[138]. When they are large enough to handle, prick the seedlings out into individual pots and grow them on in a cold frame for at least their first winter. Plant them out into their permanent positions in late spring or early summer, after the last expected frosts. The seed can also be sown outdoors in situ during the spring. Basal cuttings in spring[1]. Harvest the shoots when they are about 10 - 15cm long with plenty of underground stem. Pot them up into individual pots and keep them in light shade in a cold frame or greenhouse until they are rooting well. Plant them out in the summer. Division in spring or autumn[111]. Very easy, larger clumps can be replanted direct into their permanent positions, though it is best to pot up smaller clumps and grow them on in a cold frame until they are rooting well. Plant them out in the summer or following spring." [PFAF]

Cultivation
"A very easily grown plant, it succeeds in most fertile well-drained soils[233, 271], though it prefers a moist but well-drained rich sandy loam and a neutral or alkaline soil in sun or partial shade[1, 200]. Succeeds in poor soils[17]. This species can be naturalized in finer turfs, on grassy banks and in chalk downland and heath associations[200]. Plants are hardy to at least -15oc[200]. A very variable species in the wild[271]. When established, plants can spread fairly freely and also self-sow, though they are quite easily contained by hoeing[271]. The species in this genus do not often hybridize and so seed can generally be relied upon to come true[221]. The plants are self-fertile[221]. Members of this genus are rarely if ever troubled by browsing deer or rabbits[233]. A very ornamental plant, there are some named varieties[200]." [PFAF]

Metal Tolerant: "Associated metal-tolerant species: matrix species that are associated with the related plant association with a large ecological amplitude. They are either called ‘pseudo-metallophytes’ or ‘accompanying species’ of the true metallo- phyte vegetation. These species are moderately tolerant of heavy metals in soil, but not dependent on their presence. Examples of such species which are both common and have a wide geographic distribution are: Achillea millefolium, Campanula rotundifolia...." [Batty EIP]

Synonyms
includes C. latisepala.
  • Campanula alaskana (A. Gray) W. Wight ex J.P. Anderson
  • Campanula dubia A. DC.
  • Campanula gieseckiana Vest ex Schult.
  • Campanula gieseckiana subsp. groenlandica (Berlin) Böcher
  • Campanula gieseckiana var. arctica (Lange) Böcher
  • Campanula groenlandica Berlin
  • Campanula heterodoxa Bong.
  • Campanula intercedens Witasek
  • Campanula petiolata A. DC.
  • Campanula rotundifolia subsp. groenlandica (Berlin) A. Löve & D. Löve
  • Campanula rotundifolia subsp. intercedens (Witasek) A. Löve & D. Löve
  • Campanula rotundifolia var. alaskana A. Gray
  • Campanula rotundifolia var. alpina Tuck.
  • Campanula rotundifolia var. arctica Lange
  • Campanula rotundifolia var. intercedens (Witasek) Farw.
  • Campanula rotundifolia var. lancifolia Mert. & W.D.J. Koch
  • Campanula rotundifolia var. petiolata (A. DC.) J.K. Henry
  • Campanula rotundifolia var. rotundifolia
  • Campanula rotundifolia var. velutina A. DC.
  • Campanula sacajaweana M. Peck

References


Campanula scouleri - Scouler's harebell

Campanula scouleri 3224f [1]
Campanula scouleri

Image References

"General: Perennial herb from a branching, slender rhizome; stems lax, often curved at the base, 5-40 cm tall, glabrous to inconspicuously short-hairy." [IFBC-E-flora]

Origin Status: Native [E-flora]

"Habitat / Range: Mesic to dry forests, rock outcrops and talus slopes in the lowland zone; locally common on Vancouver Island and the Gulf Islands; N to SE AK and S to N CA." [IFBC-E-flora]

Ecological Indicator Information
"A shade-intolerant, submontane to montane. Western North American forb distributed more in the Pacific than the Cordilleran region. Occurs in cool mesothermal climates on very dry to moderately dry, nitrogen-poor soils (Mor humus forms). Sporadic in open-canopy forests on strongly drained, rocky and stony, water-shedding sites; also in non-forested communities on disturbed sites. An oxylophytic species characteristic of moisture-deficient sites." (Information applies to coastal locations only) [IPBC-E-flora]

References


Campanula trachelium - Nettle-leaved bellflower 

Campanula trachelium Dzwonek pokrzywolistny 2018-07-28 Ślęża 02 [1]
Campanula trachelium

Image References

"Garden escape (bats in the belfry) persists in the old overgrown Chinatown, Coombs, Vancouver Island. (Lomer 5728 @ UBC)." [E-flora]

Food Use

"Some plants, such as Taraxacum officinale Web., Cardaria draba (L.) Desv. and Campanula trachelium L., can be consumed raw and some without any preliminary preparation other than trimming and cleaning. They are used as an appetizer, and for flavoring, as well as for snacks at picnics and in travels." Root & Shoot of C. trachelium consumed raw. [dogan,2004]

Lore

"NETTLE-LEAVED BELLFLOWER (Campanula trachelium) Once much in demand for treating sore throat and tonsilitis (Grigson). The specific name, trachelium, shows it was a remedy for inflammation of the windpipe, or trachea (Fisher). Gerard called it Throatwort, or Uvula-wort, and other names like Haskwort or Halswort (Cockayne), which is straight from the German Halskraut, where Hals means the throat, or neck, are evidence enough of the usage. Hence, too, the early name Neckwort (Storms). Earlier still, it was taken to be a wound herb, quoted as such in the Anglo-Saxon Apuleius" [DPL Watts]

References


Page last modified on 10:48 PM October 8, 2024